These are high quality satin nickel finished tuners made by TonePros with satin nickel bushings and screws. In the mid 2010s Gibson USA installed TonePros/Kluson tuners on their Les Paul Standards as an upgrade to the usual tuners sourced from PingWorks. These are made in conjunction with Kluson and say "Kluson Deluxe" on the casing. They turn very smoothly and steadily and are a bit more accurate with a 15:1 ratio instead of the usual 14:1. The plastic tips are less green and more of a cream hue than what's often installed on Gibsons. The satin nickel finish is great if you have heavily oxidized nickel hardware and want to match

These are vintage style non-collared tuners meant for guitars with push-in bushings. They won't work with modern 10mm peghead holes (the type with a hex nut, washer showing) unless you use conversion bushings.

The twelve mounting screws are #2 size (most typical)